Ph.D. – Physiology, 1980 – Washington State University, Pullman, WA
M.S. – Physical Education, 1973 – Wisconsin State University, La Crosse, WI
B.S. – Physical Education, 1970 – Wisconsin State University, La Crosse, WI

American College of Sports Medicine, fellow
American Physiological Society, full member

Dr. Klug was recognized for his achievements by the American College of Sports Medicine with the National New Investigator Award. He was a University of Colorado Teaching Excellence Award Finalist and the recipient of graduate research fellowships from the University of Texas Health Science Center in Dallas and the Alexander Von Humboldt Foundation for study at the University of Konstanz in Germany. Klug received grants from the American Heart Association, The National Institutes of Health, Hedco Foundation, the Beckman Foundation, and the Medical Research Foundation of Oregon to support his research on the etiology of muscle fatigue at the cellular level. His research has been published in journals such as the Exercise and Sport Science Reviews, Journal of Biological Chemistry, and Journal of Applied Physiology. More recently, he has been active in curriculum and instruction revision in the areas of physiology and exercise physiology, focusing on how these disciplines prepare students with health-science career goals. To this end, he received a grant from the National Science Foundation under the title ” An Activity-Based Model for Teaching Critical Scientific Concepts”